The movie, officially titled Captain America: The Winter Soldier, is based heavily on the Winter Soldier story by Ed Brubaker.
"I can say in general that there's sort of a darker, edgier sensibility at work [in Brubaker's storyline] that we found appealing," Russo told The Huffington Post, "and that is going to find its way into Captain [America] in the modern day."
Brubaker's Winter Soldier sees Captain America's (Chris Evans in the film) dead best friend Bucky (Sebastian Stan) return as a brainwashed Russian assassin.
It has been confirmed that Stan will reprise his role in the sequel, while Anthony Mackie will join the cast.
Russo, who will co-direct alongside his brother Joe, went on to suggest that Captain America's alter ego Steve Rogers is still developing as a character, and will be shown to have changed significantly following the events of Captain America: The First Avenger.
"We're trying to grow him as a character, and certainly he's come a long way from where he started in pre-World War 2 to where he is in modern-day America.
"So the character has room for growth because of that huge journey that he's been on."
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